Evolution of Fairness in the Not Quite Ultimatum Game

The Ultimatum Game (UG) is an economic game where two players (proposer and responder) decide how to split a certain amount of money. While traditional economic theories based on rational decision making predict that the proposer should make a minimal offer and the responder should accept it, human subjects tend to behave more fairly in UG. Previous studies suggested that extra information such as reputation, empathy, or spatial structure is needed for fairness to evolve in UG. Here we show that fairness can evolve without additional information if players make decisions probabilistically and may continue interactions when the offer is rejected, which we call the Not Quite Ultimatum Game (NQUG). Evolutionary simulations of NQUG showed that the probabilistic decision making contributes to the increase of proposers' offer amounts to avoid rejection, while the repetition of the game works to responders' advantage because they can wait until a good offer comes. These simple extensions greatly promote evolution of fairness in both proposers' offers and responders' acceptance thresholds.Comment: 14 pages, 3 figure

Evolution of Human-like Social Grooming Strategies regarding Richness and Group Size

Human beings tend to cooperate with close friends, therefore they have to construct strong social relationships to recieve cooperation from others. Therefore they should have acquired their strategies of social relationship construction through an evolutionary process. The behavior of social relationship construction is know as "social grooming." In this paper, we show that there are four classes including a human-like strategy in evolutionary dynamics of social grooming strategies based on an evolutionary game simulation. Social relationship strengths (as measured by frequency of social grooming) often show a much skewed distribution (a power law distribution). It may be due to time costs constraints on social grooming, because the costs are too large to ignore for having many strong social relationships. Evolution of humans' strategies of construction of social relationships may explain the origin of human intelligence based on a social brain hypothesis. We constructed an individual-based model to explore the evolutionary dynamics of social grooming strategies. The model is based on behavior to win over others by strengthening social relationships with cooperators. The results of evolutionary simulations show the four classes of evolutionary dynamics. The results depend on total resources and the ratio of each cooperator's resource to the number of cooperators. One of the four classes is similar to a human strategy, i.e. the strategies based on the Yule--Simon process of power law.Comment: 21 pages, 10 figure

Network flow of mobile agents enhances the evolution of cooperation

We study the effect of contingent movement on the persistence of cooperation on complex networks with empty nodes. Each agent plays Prisoner's Dilemma game with its neighbors and then it either updates the strategy depending on the payoff difference with neighbors or it moves to another empty node if not satisfied with its own payoff. If no neighboring node is empty, each agent stays at the same site. By extensive evolutionary simulations, we show that the medium density of agents enhances cooperation where the network flow of mobile agents is also medium. Moreover, if the movements of agents are more frequent than the strategy updating, cooperation is further promoted. In scale-free networks, the optimal density for cooperation is lower than other networks because agents get stuck at hubs. Our study suggests that keeping a smooth network flow is significant for the persistence of cooperation in ever-changing societies.Comment: 7 pages, 5 figure

How mutation alters fitness of cooperation in networked evolutionary games

Cooperation is ubiquitous in every level of living organisms. It is known that spatial (network) structure is a viable mechanism for cooperation to evolve. Until recently, it has been difficult to predict whether cooperation can evolve at a network (population) level. To address this problem, Pinheiro et al. proposed a numerical metric, called Average Gradient of Selection (AGoS) in 2012. AGoS can characterize and forecast the evolutionary fate of cooperation at a population level. However, stochastic mutation of strategies was not considered in the analysis of AGoS. Here we analyzed the evolution of cooperation using AGoS where mutation may occur to strategies of individuals in networks. Our analyses revealed that mutation always has a negative effect on the evolution of cooperation regardless of the fraction of cooperators and network structures. Moreover, we found that mutation affects the fitness of cooperation differently on different social network structures.Comment: 6 pages, 5 figure

Transitions between homophilic and heterophilic modes of cooperation

Cooperation is ubiquitous in biological and social systems. Previous studies revealed that a preference toward similar appearance promotes cooperation, a phenomenon called tag-mediated cooperation or communitarian cooperation. This effect is enhanced when a spatial structure is incorporated, because space allows agents sharing an identical tag to regroup to form locally cooperative clusters. In spatially distributed settings, one can also consider migration of organisms, which has a potential to further promote evolution of cooperation by facilitating spatial clustering. However, it has not yet been considered in spatial tag-mediated cooperation models. Here we show, using computer simulations of a spatial model of evolutionary games with organismal migration, that tag-based segregation and homophilic cooperation arise for a wide range of parameters. In the meantime, our results also show another evolutionarily stable outcome, where a high level of heterophilic cooperation is maintained in spatially well-mixed patterns. We found that these two different forms of tag-mediated cooperation appear alternately as the parameter for temptation to defect is increased.Comment: 16 pages, 7 figure